Cause Puccinia coronata, a fungus that overwinters on infected plants. Tall fescue, perennial ryegrass, bluegrass, and orchardgrass are all susceptible. The alternate host is buckthorn (Rhamnus spp.). Its importance varies considerably with its proximity to the grass host. Primary inoculum is urediniospores; secondary inoculum (urediniospores) is produced in initial pustules and disseminated by wind. Teliospores are two-celled and are produced in late summer.