Cause Erysiphe calocladophora, Erysiphe penicillata and Phyllactinia alnicola, (formerly Microsphaera sp.), fungi that have been reported from Oregon and Washington. The conidia are readily wind disseminated. Chasmothecia develop within the external hyphae and form appendages late in their development. This fungus is favored by conditions that produce high humidity but dry leaves.