Cause The cereal cyst nematodes, Heterodera avenae and H. filipjevi, occur in at least seven Western States including Idaho, Washington, Oregon, and California. This nematode has a wide host range in the Gramineae including oats, wheat, barley, rye, triticale and ryegrass. Susceptible spring grains are barley, oats, wheat, rye, and triticale. Spring varieties are more severely affected than fall varieties because the nematode attacks in spring when plants are small.