See:
Lily (Lilium spp.) - Nematode, Root-lesion
Notes Several Pratylenchus species have been reported on chrysanthemum as most harmful including P. penetrans P. dianthus and P. kumamotoensis. These nematodes enter into roots and cause root lesions resulting in stunting and yellowing of plant shoots.
Reference Kawanobe, M., Toyota, K., Uchihara, H. and Takae, M. 2019. Developing a real-time PCR diagnostic method for a potential threat to chrysanthemum, Paratylenchus dianthus. Journal of Nematology, 51:e2019-43.